Dr. Franklin Carrero-Martínez 
Adjunct Professor (Assistant Professor of Dept. of Biology, UPR Mayaguez)
Lab B148, B-167, B170, B175, Office B-204 (Dept. of Biology, UPR Mayaguez)
franklin.carrero@upr.edu
787-832-4040 x2405 x3923
Interests: Molecular, genetic and structural influences on the development of the neuromuscular junction in Drosophila melanogaster.
The long-term goal of my research program is motivated by several issues in cellular and molecular neuroscience: The understanding of molecular mechanisms underlying development, regulation and function of synapses. In pursuit of this goal, my research program is currently focused on the underlying signaling mechanisms involved in synaptic partner-induced postsynaptic filopodial clustering and the recruitment of cytoplasmic proteins to this subcellular space.
For centuries, a key question of interest to many has been how a synapse develops. Historically, muscles were thought to be passive structures that merely provided a substrate for the active axonal growth cone. However, we now know that faithful formation of synapses results from the precise spatiotemporal interactions of the molecular components responsible for target guidance and recognition, cytoskeletal remodeling and subsynaptic specialization.
This project investigates the in vivo cellular dynamics that are suspected to have major impacts on the process of synaptogenesis. Using the Drosophila neuromuscular junction as a model, my research attempts to gain a better understanding of the three-dimensional interactions in vivo between synaptic partners, which are currently unknown.
Available high-resolution bio-imaging techniques (GFP- and RFP-based bio-probes together with time-lapse confocal microscopy) allows me to perform in vivo analysis at high cellular resolutions using live, undissected organisms.This intact embryo context preserves the spatial three-dimensionality of those molecular and physical factors playing a role in axon guidance, target recognition and synaptogenesis. It also facilitates the study of mechanisms involved in brain organization within their native environment.
GFP (green fluorescent protein) and RFP (red fluorescent protein) tagged proteins are being visualized using different acquisition techniques. Depending on the purpose of the experiment, 2D time-lapse (high temporal resolution), 3D (high spatial resolution) or 4D (3D time-lapse; a hybrid level resolution) image stacks are acquired using inverted microscopes with either laser scanning or spinning-disc confocal modules. The image stacks are then analyzed using various 3D reconstruction and deconvolution software packages. In order to facilitate the organization of this site and to facilitate discussion of this highly specific and complicated developmental event, I have divided this part of my research in three main areas: (1) Myopodia (post-synaptic filopodia); (2) Neuromuscular synaptogenesis (Muscle Innervation); (3) Muscle Contraction. In summary, this project investigates the in vivo cellular dynamics that are suspected to have major impacts of the process of synaptogenesis. The results are anticipated to enhance our hitherto underappreciated cellular dynamics that surround, or perhaps even dictate the process of synaptogenesis.
Selected Publications:
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Carrero-Martínez, F. A., Furrer, M.-P., Suzuki, E. and Chiba, A. “Drosophila Synaptogenesis requires filopodial clustering and postsynaptic Dlg/PSD-95”. In preparation.
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Siechen, S., Yang, S., Sun, J., Carrero-Martínez, F. A., Saif, T. and Chiba, A. (2009). “Mechanical force contributes to assembly and plasticity at presynaptic terminal”. Submitted Proceedings of the National Academy of Science of the USA.
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Carrero-Martínez, F. A. and Chiba, A. (2008). “Cell Adhesion Molecules at the Drosophila Neuromuscular Junction”. Introductory chapter for the “Cell Adhesion Molecules in Synaptogenesis” book by editors Michael Hortsch and Hisashi Umemori. Springer Publishing.
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Angenstein, F., Evans, A. M., Ling, S. C., Settlage, R. E., Ficarro, S., Carrero-Martínez, F. A., Shabanowitz, J. and Greenough, W. T. (2005). Proteomic characterization of mRNP-complexes bound to nontranslated or translated polyA-mRNAs in the rat cerebral cortex. Journal of Biological Chemistry 280 (8): 6496 - 503.
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Chiba, A., Carrero-Martínez, F. A. and Kamiyama, D. (2003). Behold, the synapse is born. Saibo Kogaku (Cell Technology) 22 (7): 726 - 729. [In Japanese].
Selected abstracts/presentations:
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Carrero-Martinez, F. and Siritunga, D. BYOC: Research in the undergraduate teaching laboratory. 2009. 3rd Annual Understanding Interventions that Encourage Minorities to Pursue Science Careers Conference, Bethesda, MD.
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Siechen, S., Sun, J., Yang, S. Carrero-Martinez, F., Saif, T. and Chiba, A. Mechanical Force Locally Adjust Accumulation of Synaptic Proteins at Axon Terminal. 2006. 46th American Society for Cell Biology Meeting, San Diego, CA.
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Synaptogenesis Requires Postsynaptic PSD-95/Dlg. Receptors, Channels and Synapses. 2006. Cold Spring Harbor Laboratory. Cold Spring Harbor, NY.
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Essential role of Postsynaptic PSD-95/Dlg and its Collaboration with Myopodial Cluster at the onset of Glutamatergic Neuromuscular Synaptogenesis in Drosophila. Neurobiology of Drosophila. 2005. Cold Spring Harbor Laboratory. Cold Spring Harbor, NY.
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Postsynaptic Filopodia (Myopodia) and Subsynaptic Reticulum at the Drosophila Neuromuscular Junction: A Potential Link. 7th Annual Japan Drosophila Research Conference. 2005. Carrero-Martínez, Franklin Awaji Island, Japan.
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In vivo Neuromuscular Synaptogenesis: Interactions, Contacts and a Role for Dlg/PSD95. Imaging Neurons and Neural Activity: New Methods, New Results. 2005. Cold Spring Harbor Laboratory. Cold Spring Harbor, NY.
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3D in vivo Analysis of Drosophila Neuromuscular Synaptogenesis. 6th International Brain Research Organization World Congress. 2003. Prague, Czech Republic & 32nd Society for Neuroscience Meeting. 2002. Orlando, FL.
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Tridimensional Analysis of Neuromuscular Synaptogenesis: Role of Dlg. Midwest Fly Meeting. 2003. Allerton, IL.
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The Fragile X Mental Retardation Protein (FMRP) is an in vitro Substrate for Protein Kinase C and Protein Kinase A”. 30th Society for Neuroscience Meeting. 1999. Miami, FL.
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Proximal-Distal Effects and Polarizing Activity in the Hind Limb Regeneration of Xenopus laevii. International Symposium on Metamorphosis and Regeneration: Keys to Tissue Regeneration. 1997. Indianapolis, IN.
